There is plenty of evidence for what scientists refer to as primary emotions – happiness and fear, for example – in animals. But empirical evidence for secondary emotions like jealousy, pride, and guilt, is extremely rare in the animal cognition literature. The argument usually given for this lack of evidence is that such secondary emotions seem to require a level of cognitive sophistication, particularly when it comes to self-awareness or self-consciousness, that may not exist in non-human animals. In other words, guilt is complicated.
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