Wikipedia is a great example of the difficulties in maintaining a positive culture over time in the face of growth. What marks them out especially is the way they recognise the problem, quantify it, and are trying to fix it.
Wikipedia has had a decline in new active editors that began in 2007. "While just as many productive contributors enter the project today as in 2006, they are entering an environment that is increasingly challenging, critical, and/or hostile to their work."
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